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Image Licensed under GNU == Defining Mutations ==* M253, M307.2/P203.2, M450/S109, P30, P40, L64, L75, L80, L81, L118, L121/S62, L123, L124/S64, L125/S65, L157.1, L186, L187== Origin == Origins. According to a study published in 2010, I-M253 originated between 3,170 and 5,000 years ago, in Chalcolithic Europe. ... A 2014 study in Hungary uncovered remains of two individuals from the Linear ...